Biological soil crusts (BSCs) are complex matrices of soil particles, mosses, lichens, and cyanobacteria that prevent erosion and influence water and energy balances, soil fertility, and vascular plant germination. The processes that form BSCs, the factors that control their distribution, and the ecosystem feedbacks that they sustain are poorly understood. This dissertation employed a novel interdisciplinary approach to address those research unknowns through investigations of the micromorphological structure, soil-geomorphic relationships, and biogeochemical feedbacks of BSCs in the Mojave Desert.